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2021 Update: Lilium fire and flowering

In summer 2021, Team Echinacea searched for and mapped 708 flowering Prairie lily (Lilium philadelphicum) plants across 6 patches of remnant prairie in our study area in western Minnesota. Unlike many prairie species that flower vigorously after fire, these self-incompatible lilies flower two years after dormant season fires. We are investigating how fire influences lily flowering density and reproduction. We harvested seed pods from 80 individuals across 5 patches to quantify seed set.

  • Start year: 2021
  • Location: Remnant patches of prairie in and around Solem Township, MN
  • Overlaps with: NA
  • Data collected: demographic data: ~aiisummer2021/otherSpp/Lilium_philadelphicum/output/lilium2021Data.csv
  • Samples or specimens collected: Pods/seed collected during summer 2021 currently reside in Jared’s office. These seeds need to be inventoried, cleaned, counted, and scored for seed set.
  • Products: Stay tuned!

You can read more about the Lilium fire and flowering project, as well as links to prior flog entries about this experiment, on the background page for this experiment.

2021 Update: Flowering phenology in the remnants

In 2021, we collected data on the timing of flowering for 1692 flowering plants (2292 flowering heads) in 32 remnant populations. The plants ranged from having 1 to 18 flowering heads. The earliest bloomers initiated flowering on June 18, and the latest head initiated flowering on August 1. We identify each plant with a numbered tag affixed to the base and give each head a colored twist tie, so that each head has a unique tag/twist-tie combination, or “head ID”, under which we store all phenology data. We monitor the flowering status of all flowering plants in the remnants, visiting at least once every three days (usually every two days) until all heads are done flowering to obtain start and end dates of flowering.

Linking this detailed phenology data with information about seed production will help us understand how the timing of reproduction influences pollination and reproductive fitness. Additionally, we are excited to investigate whether fire synchronizes flowering in remnant populations. Eight of the populations in which we collected phenology data were burned during spring 2021.

Start year: 1996

Location: Roadsides, railroad rights of way, and nature preserves in and around Solem Township, MN

Overlaps withreproductive fitness in remnants

Data/materials collected: We managed the data in the R project ‘aiisummer2021′ and will add the records to the database of previous years’ remnant phenology records, which is located here: https://echinaceaproject.org/datasets/remnant-phen/. The 2021 phenology data set needs to be cleaned and prepared for integration with phenology data from previous years and is still located in the aiisummer2021 repo.

Products: Stay tuned!

You can find more information about phenology in the remnants and links to previous flog posts regarding this experiment at the background page for the experiment.

Updating and maintaining prescribed fire equipment

With a busy burn season ahead of us, I took some time in late October to maintain the equipment we use for prescribed burns. During a hectic spring 2021 burn season, we encountered several issues. A handful of 5-gallon backpack pumps were not functioning well and one of our drip torches was constantly acting up (especially while using the leftover “spicy” fuel mix).

During spring 2021, I noticed two of our 5-gallon backpack pumps had cracked pump cylinders and would need to be replaced. I tried applying plumber’s epoxy to the cracked cylinders last spring but the pressure created by the pump inevitably caused this epoxy to fail. Over the summer, we purchased replacement outer cylinders for the backpack pump assemblies. I rebuilt the pump assemblies and installed these cylinders last week. The pumps seems to be working properly now. We should be very diligent about making sure all water has been expelled from tanks and pump assemblies prior to winter storage to avoid cracked cylinders.

We also purchased four harnesses and two pairs of padded straps for our 5-gallon pumps. These provide a welcome upgrade over the notoriously uncomfortable thin straps that come with the backpack pumps. We now should have 6 fully functional (and comfortable!) backpack pumps for burning.

Rebuilding the troublesome drip torch was a priority for me. After inspecting the torch, I realized one if not the problem was a badly deteriorating collar gasket (the big O-ring). I replaced this gasket along with the smaller O-ring on the discharge plug. Pro-tip: Forestry Suppliers sells discharge plug O-rings for drip torches but the ones they sell are too large. I found the #9 O-rings (5/8 in. outer diameter x 7/16 in. inner diameter x 3/32 in. wall) that are readily available at just about any hardware store work much better. I also replaced the breather valve assembly on this drip torch.

This brings up a more general point about the importance of maintaining burn equipment. At the beginning of the burn season we should:

  • Check to make sure all 5-gallon backpack pumps are functioning properly
    • Check to make sure gasket is intact and installed in top lid
    • Inspect pump cylinder for cracks
    • Inspect nozzle and make sure it is clear of debris
    • Install paper clip used to clear nozzle obstructions
    • Ensure nozzle is set on adapter with two holes (single hole adapter not very effective or water-efficient for extinguishing grass fires)
    • Fill each tank with water and test pump
  • Check drip torches
    • Inspect integrity of collar gasket, replace immediately if damaged
    • Inspect integrity of discharge plug O-ring, replace if damaged
    • Check for obstructions or debris that could impede flow of fuel
  • Rakes and swatters
    • Locate metal rakes and swatters
    • Check integrity of the swatters (these can melt and deteriorate making them ineffective)
    • Note any tools that need replacing
  • Kestrel
    • Locate Kestrel 3500FW and test unit/check battery
    • Locate or purchase additional (new) battery

At the end of the season:

  • 5-gallon backpack pumps
    • Ensure all water and excess moisture has been expelled from 5-gallon pumps
    • Inspect integrity of gasket on tank lid
  • Drip torch
    • If empty, use paper towel to clean interior and remove debris
    • Check integrity of collar gasket and discharge plug O-rings
  • Return rakes and swatters to G3
  • Hang Kestrel 3500FW from wooden dowel above shelf between main room and bathroom in Hjelm

Wrapping up the field season

It has been a hectic couple weeks for the Echinacea Project. Last week we braved fog, wind, and rain to wrap up the 2021 field season. Mia and Alex drove up from Chicago to help finish planting and pick up flags in the remnants.

On Thursday and Friday, Mia, Alex, and I sowed Echinacea seed in the 76 transects that comprise our seed addition experiment. This experiment will help us quantify the effects of fire and other environmental variables on seedling emergence and survival.

The planting went smoothly, we averaging approximately 2 minutes per transect. Prior to sowing seeds, we located transects and found nails designating the start and end of focal 1-m segments. We laid a meter stick on the ground stretching from start nail to end nail. We then spread seeds evenly between the 5 and 95 cm marks of the meter stick in line with the transect. Once empty, each seed envelope was given one last flick to dislodge any stubborn achenes and we gently tapped the meter stick against the ground to ensure any achenes that landed on blades of grass or other vegetation were not catapulted across the prairie when we picked up the meter stick.

Barring a shift in the weather that brings more favorable conditions for burning, Team Echinacea has moved indoors for the winter.

Establishing seed addition transects

As part of the 2020 NSF grant to study fire effects on plant reproduction and population dynamics, we are implementing a seed addition experiment in numerous remnants. From previous studies, we know that fire can improve recruitment which is important for population growth. However, our previous observations of recruitment in remnants conflate the amount of seed entering the seed bank and the seedlings emerging from the seed bank. The goal of this seed addition experiment is to help us directly quantify the effects of fire on seedling emergence and early seedling fitness. We will use these data to parameterize demographic models for Echinacea.

For the seed addition experiment, we established 76 transects distributed across 32 prairie remnants with Echinacea. Transect locations were determined by generating an ordered list of random points (random integers corresponding with MN state plane coordinate system) within each remnant and selecting the first 2-4 random points that were located within ~5m of an adult Echinacea but avoided dense patches of flowering plants where we may have difficulty distinguishing experimental seedlings from natural recruits. Each transect originating at a random point is 4-m long and contains four 1-m segments. Most transects extend North from the random point but some extend East (in sites where North-South transects may span an entire ditch). One segment per transect was chosen at random to be planted in fall 2021 and one transect chosen at random to be planted during fall 2022. The study includes 9 sites burned during spring 2021 as well as 7 sites slated to burn during spring 2022.

Over the past week, I have been using a GPS unit to stake the transect locations. I marked the start and end point of each transect using a blue pin flag and installed nails at 1-m intervals using measuring tape (this will facilitate the use of meter sticks and/or measuring tapes in the field). We used 4 inch galvanized common nails. Pro tip: these nails were considerably less expensive at Fleet Farm than other vendors. I plan to finish setting up transects either this afternoon or tomorrow afternoon. Our goal is to begin sowing seeds later this week.

Days growing shorter and colder

After a balmy stretch of weather (at least balmy for Minnesota in October), colder temperatures and shorter days have descended upon western MN. We experienced a killing frost Saturday (Oct 16) morning. Dwight and Jean reported that their tomatoes and other frost-sensitive garden plants are done for the year. I also awoke to frost this morning after a cold and rainy Wednesday (Oct 20).

During two of the next three nights, overnight lows are predicted to dip below 30 F. The forecast is also calling for a chance of snow (mixed with rain) Sunday morning. We had hoped for a dry, sunny stretch of weather this fall favorable for burning but the next week looks iffy.

The end of summer

As summer’s vibrant greens give way to fall’s golden glow, Team Echinacea remains hard at work in western MN. A skeleton crew is diligently wrapping up the field season. Our most important task is harvesting seed from study species so that we can quantify fire effects on plant reproduction in remnant prairies. Here is a brief update on progress for our focal species:

Echinacea angustifolia harvest: 383/383 plants harvested

Andropogon gerardii harvest: 370/370 plots measured and harvested

Liatris aspera harvest: 202/231 plants harvested

Lilium philadelphicum harvest: 79/80 plants harvested

Asclepias viridiflora: all plants harvested (~30, Jared forgot to check harvest data sheet…)

In addition to wrapping up the harvest, we are beginning to make preparations for fall burns and getting materials organized to implement a seed addition experiment designed to assess fire effects on seedling emergence and survival.

keeping the bees busy

Today marked the beginning of our third week of the field season. The team began the day with phenology measurements in the remnants. Today we measured 509 flowering Echinacea heads! This is roughly 1/4 to 1/3 of the heads we are monitoring. We expect flowering will peak next week. Stay tuned for more information! During the afternoon, the team worked on independent projects and continued assessing Hesperostipa reproduction in remnants. Emma and Amy measured a plot at landfill east with over 100 culms… oofta!

We are staying busy, and so are the bees!

things are about to get busy

On Saturday morning, Amy and I each visited a handful of remnants to check on Echinacea flowering status. Amy found a few “rays spreading” plants but none shedding pollen. I found 7 heads shedding pollen (2 heads at eri, 3 heads at ri, 1 head at lc, and 1 head at ness). A couple heads were on their second day of flowering making June 18 the first day of flowering in our study sites. Notably, several big bluestem have started bolting including one at ri with a culm taller than my waist. Oofta.

Over the past few days, I also spent some time searching for wood lilies (Lilium philadelphicum), also known as prairie lilies or western red lilies. When flowering, these jaw-dropping beauties are hard to miss! The vivid red-orange tepals stand out, bright flames flickering amidst a drab sea of greens and browns. I am conducting a preliminary study of lily reproduction this summer in hopes of examining how these charismatic wildflowers respond to fire. We are still early, most lily buds remain tight and green, but I have developed an eye for the buds. My plan is to visit a number of sites next week. I might be slightly obsessed with lilies…

And so it begins…

For many months I have been eagerly awaiting summer in the prairie. After a long winter and a busy spring, I found visiting local prairies during the past couple weeks incredibly restorative. Each step is accompanied by a familiar crunch as new “green friends” come into view. Their distinctive color palettes fuse into a vivid mosaic. As the summer progresses, I know incredible hues of green, yellow, white, purple, and orange will wax and wane producing a shifting mosaic of colors that simply enthralls the senses. I know what is coming, yet I can hardly contain my excitement.

Welcome Team Echinacea 2021! Here’s to a safe, productive, and illuminating summer of science filled with kaleidoscopic prairie sunsets and a menagerie of new friends, green and otherwise.